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The players' Keystones work with any Mega Stone they want, so I assume they're interchangeable in the games as well.So you can borrow keystones, well I guess that's important or else how would people be able to mega evolve? I mean it's pretty obvious, but that might have to do with the plot at one time.
They do, actually. When Ash and Bonnie were asking if Pikachu and Dedenne could Mega Evolve, Sycamore said it wouldn't happen with a non-final evolution like Pikachu, but didn't rule it out for Dedenne. 'Course, a Dedennite would need to exist for that to happen.Also it seems they don't know that mega evolution happens at the last stage of a Pokemon